epigean

That lives on or near the surface of the land.

For other Leptodirini, the contraction of the life cycle is less radical, being intermediate between this extreme and a epigean life cycle.

Any organism that lives on or near the surface of the land.

We also studied the female internal system for a species in the subgenus Aphaenops (Cerbaphaenops), and two species belonging to two other genera, one endogean (Geotrechus) and one epigean (Trechus).